{"id":329403,"date":"2026-02-11T11:31:02","date_gmt":"2026-02-11T10:31:02","guid":{"rendered":"https:\/\/isem-evolution.fr\/membre\/ryo\/"},"modified":"2026-02-11T11:32:21","modified_gmt":"2026-02-11T10:32:21","slug":"ryo","status":"publish","type":"membre","link":"https:\/\/isem-evolution.fr\/en\/membre\/ryo\/","title":{"rendered":"Ryo"},"content":{"rendered":"","protected":false},"featured_media":329402,"template":"","meta":{"_acf_changed":false,"site-sidebar-layout":"default","site-content-layout":"","ast-site-content-layout":"default","site-content-style":"default","site-sidebar-style":"default","ast-global-header-display":"","ast-banner-title-visibility":"","ast-main-header-display":"","ast-hfb-above-header-display":"","ast-hfb-below-header-display":"","ast-hfb-mobile-header-display":"","site-post-title":"","ast-breadcrumbs-content":"","ast-featured-img":"","footer-sml-layout":"","ast-disable-related-posts":"","theme-transparent-header-meta":"default","adv-header-id-meta":"","stick-header-meta":"","header-above-stick-meta":"","header-main-stick-meta":"","header-below-stick-meta":"","astra-migrate-meta-layouts":"set","ast-page-background-enabled":"default","ast-page-background-meta":{"desktop":{"background-color":"","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"tablet":{"background-color":"","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"mobile":{"background-color":"","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""}},"ast-content-background-meta":{"desktop":{"background-color":"var(--ast-global-color-5)","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"tablet":{"background-color":"var(--ast-global-color-5)","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"mobile":{"background-color":"var(--ast-global-color-5)","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""}}},"categories_membres":[46],"class_list":["post-329403","membre","type-membre","status-publish","has-post-thumbnail","hentry","categories_membres-phdstudents-en"],"acf":[],"yoast_head":"\nAlexis Ryo - ISEM<\/title>\n<meta name=\"description\" content=\"My research lies within the field of evolutionary biology and focuses on the evolution of reproductive systems in plants. My PhD thesis aims to document and compare the intensity of sexual dimorphism in ten dioecious plant species belonging to the genus Silene, in order to better understand the proximate and ultimate mechanisms that promote, or conversely constrain, the emergence of trait differences between male and female individuals. This comparative approach is based on the analysis of vegetative traits (number of leaves, dry biomass) and reproductive traits (number and size of flowers, olfactory emissions, sugar content in nectar). The thesis is structured around two central questions: 1- the impact of the age of dioecy on the intensity of sexual dimorphism 2- the impact of pollination mode (entomophilous vs anemophilous) on the expression of sexual dimorphism\" \/>\n<meta name=\"robots\" content=\"index, follow, max-snippet:-1, max-image-preview:large, max-video-preview:-1\" \/>\n<link rel=\"canonical\" href=\"https:\/\/isem-evolution.fr\/en\/membre\/ryo\/\" \/>\n<meta property=\"og:locale\" content=\"en_US\" \/>\n<meta property=\"og:type\" content=\"article\" \/>\n<meta property=\"og:title\" content=\"Alexis Ryo - ISEM\" \/>\n<meta property=\"og:description\" content=\"My research lies within the field of evolutionary biology and focuses on the evolution of reproductive systems in plants. My PhD thesis aims to document and compare the intensity of sexual dimorphism in ten dioecious plant species belonging to the genus Silene, in order to better understand the proximate and ultimate mechanisms that promote, or conversely constrain, the emergence of trait differences between male and female individuals. This comparative approach is based on the analysis of vegetative traits (number of leaves, dry biomass) and reproductive traits (number and size of flowers, olfactory emissions, sugar content in nectar). The thesis is structured around two central questions: 1- the impact of the age of dioecy on the intensity of sexual dimorphism 2- the impact of pollination mode (entomophilous vs anemophilous) on the expression of sexual dimorphism\" \/>\n<meta property=\"og:url\" content=\"https:\/\/isem-evolution.fr\/en\/membre\/ryo\/\" \/>\n<meta property=\"og:site_name\" content=\"ISEM\" \/>\n<meta property=\"article:modified_time\" content=\"2026-02-11T10:32:21+00:00\" \/>\n<meta property=\"og:image\" content=\"https:\/\/isem-evolution.fr\/wp-content\/uploads\/2026\/02\/RYO_Alexis_ID-e1770806906394.jpg\" \/>\n\t<meta property=\"og:image:width\" content=\"653\" \/>\n\t<meta property=\"og:image:height\" content=\"900\" \/>\n\t<meta property=\"og:image:type\" content=\"image\/jpeg\" \/>\n<meta name=\"twitter:card\" content=\"summary_large_image\" \/>\n<meta name=\"twitter:label1\" content=\"Est. reading time\" \/>\n\t<meta name=\"twitter:data1\" content=\"1 minute\" \/>\n<script type=\"application\/ld+json\" class=\"yoast-schema-graph\">{\"@context\":\"https:\\\/\\\/schema.org\",\"@graph\":[{\"@type\":\"WebPage\",\"@id\":\"https:\\\/\\\/isem-evolution.fr\\\/en\\\/membre\\\/ryo\\\/\",\"url\":\"https:\\\/\\\/isem-evolution.fr\\\/en\\\/membre\\\/ryo\\\/\",\"name\":\"Alexis Ryo - ISEM\",\"isPartOf\":{\"@id\":\"https:\\\/\\\/isem-evolution.fr\\\/en\\\/#website\"},\"primaryImageOfPage\":{\"@id\":\"https:\\\/\\\/isem-evolution.fr\\\/en\\\/membre\\\/ryo\\\/#primaryimage\"},\"image\":{\"@id\":\"https:\\\/\\\/isem-evolution.fr\\\/en\\\/membre\\\/ryo\\\/#primaryimage\"},\"thumbnailUrl\":\"https:\\\/\\\/isem-evolution.fr\\\/wp-content\\\/uploads\\\/2026\\\/02\\\/RYO_Alexis_ID-e1770806906394.jpg\",\"datePublished\":\"2026-02-11T10:31:02+00:00\",\"dateModified\":\"2026-02-11T10:32:21+00:00\",\"description\":\"My research lies within the field of evolutionary biology and focuses on the evolution of reproductive systems in plants. My PhD thesis aims to document and compare the intensity of sexual dimorphism in ten dioecious plant species belonging to the genus Silene, in order to better understand the proximate and ultimate mechanisms that promote, or conversely constrain, the emergence of trait differences between male and female individuals. This comparative approach is based on the analysis of vegetative traits (number of leaves, dry biomass) and reproductive traits (number and size of flowers, olfactory emissions, sugar content in nectar). The thesis is structured around two central questions: 1- the impact of the age of dioecy on the intensity of sexual dimorphism 2- the impact of pollination mode (entomophilous vs anemophilous) on the expression of sexual dimorphism\",\"breadcrumb\":{\"@id\":\"https:\\\/\\\/isem-evolution.fr\\\/en\\\/membre\\\/ryo\\\/#breadcrumb\"},\"inLanguage\":\"en-US\",\"potentialAction\":[{\"@type\":\"ReadAction\",\"target\":[\"https:\\\/\\\/isem-evolution.fr\\\/en\\\/membre\\\/ryo\\\/\"]}]},{\"@type\":\"ImageObject\",\"inLanguage\":\"en-US\",\"@id\":\"https:\\\/\\\/isem-evolution.fr\\\/en\\\/membre\\\/ryo\\\/#primaryimage\",\"url\":\"https:\\\/\\\/isem-evolution.fr\\\/wp-content\\\/uploads\\\/2026\\\/02\\\/RYO_Alexis_ID-e1770806906394.jpg\",\"contentUrl\":\"https:\\\/\\\/isem-evolution.fr\\\/wp-content\\\/uploads\\\/2026\\\/02\\\/RYO_Alexis_ID-e1770806906394.jpg\",\"width\":653,\"height\":900},{\"@type\":\"BreadcrumbList\",\"@id\":\"https:\\\/\\\/isem-evolution.fr\\\/en\\\/membre\\\/ryo\\\/#breadcrumb\",\"itemListElement\":[{\"@type\":\"ListItem\",\"position\":1,\"name\":\"Accueil\",\"item\":\"https:\\\/\\\/isem-evolution.fr\\\/en\\\/\"},{\"@type\":\"ListItem\",\"position\":2,\"name\":\"Membres\",\"item\":\"https:\\\/\\\/isem-evolution.fr\\\/en\\\/membre\\\/\"},{\"@type\":\"ListItem\",\"position\":3,\"name\":\"Ryo\"}]},{\"@type\":\"WebSite\",\"@id\":\"https:\\\/\\\/isem-evolution.fr\\\/en\\\/#website\",\"url\":\"https:\\\/\\\/isem-evolution.fr\\\/en\\\/\",\"name\":\"ISEM\",\"description\":\"Institut des Sciences de l'\u00c9volution de Montpellier\",\"publisher\":{\"@id\":\"https:\\\/\\\/isem-evolution.fr\\\/en\\\/#organization\"},\"potentialAction\":[{\"@type\":\"SearchAction\",\"target\":{\"@type\":\"EntryPoint\",\"urlTemplate\":\"https:\\\/\\\/isem-evolution.fr\\\/en\\\/?s={search_term_string}\"},\"query-input\":{\"@type\":\"PropertyValueSpecification\",\"valueRequired\":true,\"valueName\":\"search_term_string\"}}],\"inLanguage\":\"en-US\"},{\"@type\":\"Organization\",\"@id\":\"https:\\\/\\\/isem-evolution.fr\\\/en\\\/#organization\",\"name\":\"Institut des Sciences de l'\u00c9volution de Montpellier\",\"url\":\"https:\\\/\\\/isem-evolution.fr\\\/en\\\/\",\"logo\":{\"@type\":\"ImageObject\",\"inLanguage\":\"en-US\",\"@id\":\"https:\\\/\\\/isem-evolution.fr\\\/en\\\/#\\\/schema\\\/logo\\\/image\\\/\",\"url\":\"https:\\\/\\\/isem-evolution.fr\\\/wp-content\\\/uploads\\\/2020\\\/10\\\/LOGO_ISEM.svg\",\"contentUrl\":\"https:\\\/\\\/isem-evolution.fr\\\/wp-content\\\/uploads\\\/2020\\\/10\\\/LOGO_ISEM.svg\",\"width\":760,\"height\":172,\"caption\":\"Institut des Sciences de l'\u00c9volution de Montpellier\"},\"image\":{\"@id\":\"https:\\\/\\\/isem-evolution.fr\\\/en\\\/#\\\/schema\\\/logo\\\/image\\\/\"}}]}<\/script>\n<!-- \/ Yoast SEO plugin. -->","yoast_head_json":{"title":"Alexis Ryo - ISEM","description":"My research lies within the field of evolutionary biology and focuses on the evolution of reproductive systems in plants. My PhD thesis aims to document and compare the intensity of sexual dimorphism in ten dioecious plant species belonging to the genus Silene, in order to better understand the proximate and ultimate mechanisms that promote, or conversely constrain, the emergence of trait differences between male and female individuals. This comparative approach is based on the analysis of vegetative traits (number of leaves, dry biomass) and reproductive traits (number and size of flowers, olfactory emissions, sugar content in nectar). The thesis is structured around two central questions: 1- the impact of the age of dioecy on the intensity of sexual dimorphism 2- the impact of pollination mode (entomophilous vs anemophilous) on the expression of sexual dimorphism","robots":{"index":"index","follow":"follow","max-snippet":"max-snippet:-1","max-image-preview":"max-image-preview:large","max-video-preview":"max-video-preview:-1"},"canonical":"https:\/\/isem-evolution.fr\/en\/membre\/ryo\/","og_locale":"en_US","og_type":"article","og_title":"Alexis Ryo - ISEM","og_description":"My research lies within the field of evolutionary biology and focuses on the evolution of reproductive systems in plants. My PhD thesis aims to document and compare the intensity of sexual dimorphism in ten dioecious plant species belonging to the genus Silene, in order to better understand the proximate and ultimate mechanisms that promote, or conversely constrain, the emergence of trait differences between male and female individuals. This comparative approach is based on the analysis of vegetative traits (number of leaves, dry biomass) and reproductive traits (number and size of flowers, olfactory emissions, sugar content in nectar). The thesis is structured around two central questions: 1- the impact of the age of dioecy on the intensity of sexual dimorphism 2- the impact of pollination mode (entomophilous vs anemophilous) on the expression of sexual dimorphism","og_url":"https:\/\/isem-evolution.fr\/en\/membre\/ryo\/","og_site_name":"ISEM","article_modified_time":"2026-02-11T10:32:21+00:00","og_image":[{"width":653,"height":900,"url":"https:\/\/isem-evolution.fr\/wp-content\/uploads\/2026\/02\/RYO_Alexis_ID-e1770806906394.jpg","type":"image\/jpeg"}],"twitter_card":"summary_large_image","twitter_misc":{"Est. reading time":"1 minute"},"schema":{"@context":"https:\/\/schema.org","@graph":[{"@type":"WebPage","@id":"https:\/\/isem-evolution.fr\/en\/membre\/ryo\/","url":"https:\/\/isem-evolution.fr\/en\/membre\/ryo\/","name":"Alexis Ryo - ISEM","isPartOf":{"@id":"https:\/\/isem-evolution.fr\/en\/#website"},"primaryImageOfPage":{"@id":"https:\/\/isem-evolution.fr\/en\/membre\/ryo\/#primaryimage"},"image":{"@id":"https:\/\/isem-evolution.fr\/en\/membre\/ryo\/#primaryimage"},"thumbnailUrl":"https:\/\/isem-evolution.fr\/wp-content\/uploads\/2026\/02\/RYO_Alexis_ID-e1770806906394.jpg","datePublished":"2026-02-11T10:31:02+00:00","dateModified":"2026-02-11T10:32:21+00:00","description":"My research lies within the field of evolutionary biology and focuses on the evolution of reproductive systems in plants. My PhD thesis aims to document and compare the intensity of sexual dimorphism in ten dioecious plant species belonging to the genus Silene, in order to better understand the proximate and ultimate mechanisms that promote, or conversely constrain, the emergence of trait differences between male and female individuals. This comparative approach is based on the analysis of vegetative traits (number of leaves, dry biomass) and reproductive traits (number and size of flowers, olfactory emissions, sugar content in nectar). The thesis is structured around two central questions: 1- the impact of the age of dioecy on the intensity of sexual dimorphism 2- the impact of pollination mode (entomophilous vs anemophilous) on the expression of sexual dimorphism","breadcrumb":{"@id":"https:\/\/isem-evolution.fr\/en\/membre\/ryo\/#breadcrumb"},"inLanguage":"en-US","potentialAction":[{"@type":"ReadAction","target":["https:\/\/isem-evolution.fr\/en\/membre\/ryo\/"]}]},{"@type":"ImageObject","inLanguage":"en-US","@id":"https:\/\/isem-evolution.fr\/en\/membre\/ryo\/#primaryimage","url":"https:\/\/isem-evolution.fr\/wp-content\/uploads\/2026\/02\/RYO_Alexis_ID-e1770806906394.jpg","contentUrl":"https:\/\/isem-evolution.fr\/wp-content\/uploads\/2026\/02\/RYO_Alexis_ID-e1770806906394.jpg","width":653,"height":900},{"@type":"BreadcrumbList","@id":"https:\/\/isem-evolution.fr\/en\/membre\/ryo\/#breadcrumb","itemListElement":[{"@type":"ListItem","position":1,"name":"Accueil","item":"https:\/\/isem-evolution.fr\/en\/"},{"@type":"ListItem","position":2,"name":"Membres","item":"https:\/\/isem-evolution.fr\/en\/membre\/"},{"@type":"ListItem","position":3,"name":"Ryo"}]},{"@type":"WebSite","@id":"https:\/\/isem-evolution.fr\/en\/#website","url":"https:\/\/isem-evolution.fr\/en\/","name":"ISEM","description":"Institut des Sciences de l'\u00c9volution de Montpellier","publisher":{"@id":"https:\/\/isem-evolution.fr\/en\/#organization"},"potentialAction":[{"@type":"SearchAction","target":{"@type":"EntryPoint","urlTemplate":"https:\/\/isem-evolution.fr\/en\/?s={search_term_string}"},"query-input":{"@type":"PropertyValueSpecification","valueRequired":true,"valueName":"search_term_string"}}],"inLanguage":"en-US"},{"@type":"Organization","@id":"https:\/\/isem-evolution.fr\/en\/#organization","name":"Institut des Sciences de l'\u00c9volution de Montpellier","url":"https:\/\/isem-evolution.fr\/en\/","logo":{"@type":"ImageObject","inLanguage":"en-US","@id":"https:\/\/isem-evolution.fr\/en\/#\/schema\/logo\/image\/","url":"https:\/\/isem-evolution.fr\/wp-content\/uploads\/2020\/10\/LOGO_ISEM.svg","contentUrl":"https:\/\/isem-evolution.fr\/wp-content\/uploads\/2020\/10\/LOGO_ISEM.svg","width":760,"height":172,"caption":"Institut des Sciences de l'\u00c9volution de Montpellier"},"image":{"@id":"https:\/\/isem-evolution.fr\/en\/#\/schema\/logo\/image\/"}}]}},"_links":{"self":[{"href":"https:\/\/isem-evolution.fr\/en\/wp-json\/wp\/v2\/membre\/329403","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/isem-evolution.fr\/en\/wp-json\/wp\/v2\/membre"}],"about":[{"href":"https:\/\/isem-evolution.fr\/en\/wp-json\/wp\/v2\/types\/membre"}],"wp:featuredmedia":[{"embeddable":true,"href":"https:\/\/isem-evolution.fr\/en\/wp-json\/wp\/v2\/media\/329402"}],"wp:attachment":[{"href":"https:\/\/isem-evolution.fr\/en\/wp-json\/wp\/v2\/media?parent=329403"}],"wp:term":[{"taxonomy":"categories_membres","embeddable":true,"href":"https:\/\/isem-evolution.fr\/en\/wp-json\/wp\/v2\/categories_membres?post=329403"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}